LONDON: Five European nations say Russian opposition leader Alexei Navalny was poisoned with a lethal toxin, and are blaming the Russian state for the attack.

The foreign ministries of the UK, France, Germany, Sweden and the Netherlands said Saturday that analysis of samples from Navalny, who died two years ago, “have conclusively confirmed the presence of epibatidine.”

It is a toxin found in poison dart frogs in South America.

The countries said that “only the Russian state had the combined means, motive and disregard for international law” to carry out the attack.